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Johana’s research interests include sexual selection and mating systems, with a particular interest in species that exhibit parental care behavior. In her research, Johana has used frogs as her study system. For her dissertation work, she focused on elucidating the parental care behavior of the smooth guardian frog of Borneo (Limnonectes palavanensis), an unusual species about which very little was known when she began her work. She is interested in understanding this species’ mating system and what behavioral and ecological factors shaped its evolution.